diff --git a/changelog.txt b/changelog.txt
index 110d1905..614e4ae1 100644
--- a/changelog.txt
+++ b/changelog.txt
@@ -39,6 +39,9 @@ OPENSEADRAGON CHANGELOG
 * Fixed: zoomTo/zoomBy ignore refPoint if immediately is true (#1184)
 * Enabled configuration of ImageLoader timeout (#1192)
 * Viewer.open() now supports an initialPage argument for sequenceMode (#1196)
+* Fixed: IIPImageServer didn't work with the latest OSD release (#1199)
+* Now clamping pixel ratio density to a minimum of 1, fixing display issues on low density devices (#1200)
+* Improved calculation for determining which level to load first (#1198)
 
 2.2.1:
 
diff --git a/src/dzitilesource.js b/src/dzitilesource.js
index 54b8cb68..443bfaca 100644
--- a/src/dzitilesource.js
+++ b/src/dzitilesource.js
@@ -140,7 +140,7 @@ $.extend( $.DziTileSource.prototype, $.TileSource.prototype, /** @lends OpenSead
 
         if (url && !options.tilesUrl) {
             options.tilesUrl = url.replace(
-                    /([^\/]+?)(\.(dzi|xml|js))?(\?[^\/]*)?\/?$/, '$1_files/');
+                    /([^\/]+?)(\.(dzi|xml|js)?(\?[^\/]*)?)?\/?$/, '$1_files/');
 
             if (url.search(/\.(dzi|xml|js)\?/) != -1) {
                 options.queryParams = url.match(/\?.*/);
diff --git a/src/openseadragon.js b/src/openseadragon.js
index 8ed6adcb..9b07d922 100644
--- a/src/openseadragon.js
+++ b/src/openseadragon.js
@@ -875,7 +875,8 @@ function OpenSeadragon( options ){
     };
 
     /**
-     * A ratio comparing the device screen's pixel density to the canvas's backing store pixel density. Defaults to 1 if canvas isn't supported by the browser.
+     * A ratio comparing the device screen's pixel density to the canvas's backing store pixel density,
+     * clamped to a minimum of 1. Defaults to 1 if canvas isn't supported by the browser.
      * @member {Number} pixelDensityRatio
      * @memberof OpenSeadragon
      */
@@ -888,7 +889,7 @@ function OpenSeadragon( options ){
                                     context.msBackingStorePixelRatio ||
                                     context.oBackingStorePixelRatio ||
                                     context.backingStorePixelRatio || 1;
-            return devicePixelRatio / backingStoreRatio;
+            return Math.max(devicePixelRatio, 1) / backingStoreRatio;
         } else {
             return 1;
         }
diff --git a/src/tiledimage.js b/src/tiledimage.js
index 0b3c0a4e..52951f2a 100644
--- a/src/tiledimage.js
+++ b/src/tiledimage.js
@@ -980,7 +980,7 @@ $.extend($.TiledImage.prototype, $.EventSource.prototype, /** @lends OpenSeadrag
             var targetZeroRatio = viewport.deltaPixelsFromPointsNoRotate(
                 this.source.getPixelRatio(
                     Math.max(
-                        this.source.getClosestLevel(viewport.containerSize) - 1,
+                        this.source.getClosestLevel(),
                         0
                     )
                 ),
diff --git a/src/tilesource.js b/src/tilesource.js
index 26c220b7..e83c4fab 100644
--- a/src/tilesource.js
+++ b/src/tilesource.js
@@ -320,25 +320,20 @@ $.TileSource.prototype = {
 
     /**
      * @function
-     * @param {Rect} rect
+     * @returns {Number} The highest level in this tile source that can be contained in a single tile.
      */
-    getClosestLevel: function( rect ) {
+    getClosestLevel: function() {
         var i,
-            tilesPerSide,
             tiles;
 
-        for( i = this.minLevel; i < this.maxLevel; i++ ){
-            tiles = this.getNumTiles( i );
-            tilesPerSide = new $.Point(
-              Math.floor( rect.x / this.getTileWidth(i) ),
-              Math.floor( rect.y / this.getTileHeight(i) )
-            );
-
-            if( tiles.x + 1 >= tilesPerSide.x && tiles.y + 1 >= tilesPerSide.y ){
+        for (i = this.minLevel + 1; i <= this.maxLevel; i++){
+            tiles = this.getNumTiles(i);
+            if (tiles.x > 1 || tiles.y > 1) {
                 break;
             }
         }
-        return Math.max( 0, i - 1 );
+
+        return i - 1;
     },
 
     /**
diff --git a/test/modules/dzitilesource.js b/test/modules/dzitilesource.js
index e7eb2114..25b0c265 100644
--- a/test/modules/dzitilesource.js
+++ b/test/modules/dzitilesource.js
@@ -36,6 +36,9 @@
         testImplicitTilesUrl(
                 '/iiipsrv?DeepZoom=/path/my.dzi', '/iiipsrv?DeepZoom=/path/my_files/',
                 'querystring in dzi url should not be ignored before slashes');
+        testImplicitTilesUrl(
+                '/fcg-bin/iipsrv.fcgi?Deepzoom=123test.tif.dzi', '/fcg-bin/iipsrv.fcgi?Deepzoom=123test.tif_files/',
+                'filename in querystring does not have to contain slash');
     });
 
 }());